In 2079, a colossal red humanoid descends from space during the opening ceremony of the world's first orbital elevator, built in the Pacific to honor peace after the Third World War. It overwhelms humanity and seizes the elevator, and a long war begins against the red flying enemies called Vacant. By summer 2099, twenty years on, humanity has built the Seventh, a humanoid seventh-generation fighter, and has pushed Vacant back to the area around the elevator. At the floating Yashima military base, Miku of the special technology research department pilots the black Seventh F/SX-02 Tenra. Stored in its computer is the personality data of Liza, a girl she was once parted from, and Miku flies to stay by her side. The film is a yuri mecha action story centered on the deep bond between the two girls.